First wire frames and UX gets developed and then Manual test gets written based on them. This manual tests are sent to the developers and  in parallel Automation test cases are written in BDD using cucumber (Based on manual tests).

As you can see Manual tests are of paramount importance and each and every feature gets tracked to BDD tests these tests are in sync with manual tests which in return are linked to the requirement.

Hope this helps. Sorry for the delay.  :-)